In 1954, Godzilla, an ancient alpha predator, is lured to Bikini Atoll in an attempt to kill him with a nuclear bomb. In 1999, Monarch scientists Ishiro Serizawa and Vivienne Graham investigate a giant skeleton unearthed by a collapsed uranium mine in the Philippines. They find two giant spores; one dormant and one hatched with a trail that leads out of the cave to the sea. In Japan, the Janjira Nuclear Power Plant experiences unusual seismic activity; supervisor Joe Brody sends his wife Sandra with other technicians into the reactor.
A tremor breaches the reactor, leaving Sandra and her team unable to escape while the plant collapses. Fifteen years later, Joe and Sandra’s son Ford, a U.S. Navy explosive ordnance disposal officer, returns from a tour of duty to his wife Elle and young son Sam in San Francisco but has to immediately depart for Japan after Joe is detained for trespassing in the Janjira quarantine zone. Joe is determined to find out the cause of the meltdown, and he persuades Ford to accompany him to their old home to retrieve vital data. They discover that the zone is not contaminated by radiation and retrieve the data but are discovered and taken to a facility in the plant’s ruins.
The facility harbours a massive, glowing chrysalis that had been feeding off the plant’s reactors for the past 15 years and emitting stronger electromagnetic pulses over time. A giant winged insect-like creature emerges and escapes, destroying the facility. Joe is severely injured and dies as Ford, Serizawa, and Graham are taken to the U.S.S. Saratoga. The incident is reported around the world like an earthquake.
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